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Figure 4.Phylogeny of Petrocephalus (23 species, 52 specimens) estimated by maximum likelihood analysis of cytochrome b nucleotide sequences. Mormyrin Gnathonemus petersii, Mormyrops nigricans and Myomyrus macrops were used as outgroups to root the tree. Numbers at internal branches are bootstrap proportions (in %) shown only for interspecific relationships and when they exceed 50%. Black-filled vertical bars to the right of the tree indicate the river basin origins of the specimens. The scale bar corresponds to 0.04 substitutions per site. Petrocephalus boboto sp. n. and Petrocephalus arnegardi sp. n. are highlighted in gray.